flowers in the upper part and female flowers in the lower part. They appear in late June to July, and by autumn, the female flowers develop into spiny cupules containing 3-7 brownish nuts that are shed during October.
The bark often has a net-shaped (retiform) pattern with deep furrows or fissures running spirally in both directions up the trunk.
The tree requires a mild climate and adequate moisture for good growth and a good nut harvest. It is sensitive to late spring and early autumn frosts, and is intolerant of lime. Under forest conditions it will tolerate moderate shade well.
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